BIO201 Anatomy & Physiology
Lecture Outline 10: Physiology of Vision
I.Vision:
A. Anatomy of The Eye
1. Accessory structures
a. palpebrae (eyelids)
b. lacrimal apparatus
c. extrinsic eye muscles
i. superior/inferior obliques
ii. lateral/medial rectus
iii. superior/inferior rectus
2. Eyeball structure
a. anterior chamber behind cornea
i. filled with aqueous humor
b. posterior chamber behind lens
i. filled with vitreous humor
c. entire eyeball covered with conjunctiva
i. thin mucous membrane
ii. continuous with palbebral conjunctiva
iii. dilation of conjunctival blood vessels cause "bloodshot" eyes
d. fibrous tunic (outer coat of eyeball wall)
i. sclera ("whites" of the eyes)
ii. cornea - transparent cover, focuses image onto lens
iii. canal of Schlemm at junction of sclera/cornea
aa. drains anterior chamber fluid (aqueous humor)
e. vascular tunic (middle coat of eyeball wall)
i. choroid
aa. dark layer (rich in melanocytes)
bb. prevents backscattering of light
ii. ciliary body
aa. contains ciliary processes and ciliary muscle
bb. suspensory ligaments attach ciliary body to lens
cc. contraction of ciliary body draws it closer to lens
iii. suspensory ligaments
aa. keeps lens stretched tight
iv. iris ("colored" part of eye)
aa. pupil allows light to enter eye
bb. circular (constrictor) and radial (dilator) muscles control pupil diameter
f. neuronal tunic (inner coat of eyeball)
i. retina
aa. photoreceptors and neuronal layers
g. lens
i. fine focuses image onto retina (onto central fovea)
ii. accomodation adjusts thickness of lens to focus light
iii. relaxation of ciliary body for distant viewing
aa. suspensory ligament tension pulls, lens flattens
iv. contraction of ciliary body for near viewing
aa. tension is absorbed by ciliary body, lens thickens
A. Physiology of Vision
1. Photoreceptor layer
a. rods
i. absorb light
ii. do not differentiate color (only shades of greys)
iii. sensitive to low light levels
iv. more present in peripheral areas of retina (outside central fovea)
b. cones
i. absorb light of particular wavelengths (red, green, or blue)
ii. less sensitive in low light levels
iii. concentrated in central fovea
2. Rhodopsin light-absorbing pigment (consists of opsin + retinal)
a. rhodopsin is embedded in many membrane "discs" inside photoreceptors)
b. opsin protein portion holds retinal molecule
c. retinal molecule (derived from vitamin-A) absorbs light energy
3. "Dark current" in photoreceptors
a. Na+ stay open in photoreceptor when NO LIGHT is being absorbed
b. constant current of Na+ into cell keeps photoreceptor MOST ACTIVE IN DARKNESS
c. active photoreceptor (in dark) continually releases inhibitory neurotransmitters onto neuronal cells in retina
4. Light reactions
a. light photon absorbed by retinal
b. opsin releases "energetic" retinal ("bleaching")
c. opsin then activates transducin molecule
d. active transducin activates phosphodiesterase enzymes
e. active phosphodiesterase enzymes cause Na+ CLOSURE
f. Na+ channel closure reduces "dark current"
g. photoreceptor releases less of the inhibitory neurotransmitters onto neuronal retinal cells
h. neuronal retinal cells fire action potentials, carried by optic nerve to brain... NOW YOU SEE!
5. Structures of Visual Pathway
a. cornea
b. lens
c. retina
d. optic nerve (cranial nerve II)
e. optic chiasma (only the lateral field of view crosses here)
f. optic tract
g. lateral geniculate of thalamus (visual relay)
h. thalamus relays to visual cortex & association areas(occipital lobe)
i. thalamus also relays to hypothalamus and midbrain
j. hypothalamus relays info to pineal via superior cervical ganglia
k. midbrain regulates visual reflexes
This is only a general outline.
There may be material that has been discussed in lecture that is not included in this outline
and there may be material on this outline that has not been discussed in lecture.
Any material discussed in lecture or listed in this outline is "fair game" for the test.